In the future, Mankind has achieved a vast starfaring civilization, but is slowly and steadily succumbing to ennui or despair, often due to defeat and subjugation by a foreign invader. Rising against the general apathy, Harlock denies defeat and leads an outlaw crew aboard his starship Arcadia to undertake daring raids against Earth's oppressors.

The oppressors are the Mazone, a race of vegetable-based women who explored Earth in the mythic past and are now back to reclaim it. Only Harlock and his mismatched crew are brave enough and capable enough to face the enemy.

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