The story begins in a futuristic domed city called Romdo,[1] built to protect its citizens after a global ecological disaster. In this utopia, humans and androids (AutoReivs[1]) coexist with each other peacefully under a total management system. A series of murders committed by berserk robots infected with the Cogito virus begins to jeopardize the delicate balance of the social order. Behind the scenes, the government is conducting secret experiments on a mysterious humanoid lifeform called Proxy, which is believed to hold the key to the survival of mankind.

Re-l (also spelled Real, pronounced "Rielle") Mayer is assigned to investigate some of the murders with her AutoReiv partner Iggy. She encounters a Cogito-infected AutoReiv and a fast and flexible monster. She later learns that the monster was a Proxy. The other central character, an immigrant named Vincent Law, is revealed as being the eponymous proxy, but his memories of his time in his native dome of Mosque are suppressed, or perhaps he suppresses them. After being hunted down, Vincent leaves Romdo, and Re-l tags along to discover the truth behind the Proxies and the domes.

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