The story of Rurouni Kenshin takes place during the early Meiji period in Japan. It tells the story of a peaceful wanderer named Himura Kenshin, formerly known as the assassin Hitokiri Battōsai. After participating during the Bakumatsu war, Kenshin wanders the countryside of Japan offering protection and aid to those in need as atonement for the murders he once committed as an assassin. When arriving in Tokyo, he meets a young woman named Kamiya Kaoru, who was in the middle of a fight with a murderer who claims to be the Hitokiri Battōsai from her swordmanship school. Kenshin decides to help her and defeats the fake Battōsai. After discovering that Kenshin is the real Battōsai, she offers him a place stay at her dojo as she notes Kenshin is a gentle person instead. Kenshin accepts and begins to establish lifelong relationships with many people, including ex-enemies, while dealing with his fair share of enemies, new and old.

After three months of living in the dojo, Kenshin discovers that his successor as assassin of the shadows, Shishio Makoto, plans to conquer Japan by destroying the Meiji Government, starting with Kyoto. Feeling that his friends may be attacked by Shishio's faction, Kenshin goes to meet Shishio alone in order to defeat him. However, many of his friends decide to travel to Kyoto to help him in his fight. He decides to accept their help and defeats Shishio in a fight, who dies in the process due to the rise in body temperature caused by his burns.

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